by ; ;
  • ISBN13:


  • ISBN10:


  • Edition: 3rd
  • Format: Hardcover
  • Copyright: 2016-01-19
  • Publisher: Sinauer Associates is an imprint of Oxford University Press

Note: Supplemental materials are not guaranteed with Rental or Used book purchases.

Purchase Benefits

  • Free Shipping Icon Free Shipping On Orders Over $35!
    Your order must be $35 or more to qualify for free economy shipping. Bulk sales, PO's, Marketplace items, eBooks and apparel do not qualify for this offer.
  • eCampus.com Logo Get Rewarded for Ordering Your Textbooks! Enroll Now
  • Buyback Icon We Buy This Book Back!
    In-Store Credit: $4.37
    Check/Direct Deposit: $4.16
    PayPal: $4.16
List Price: $234.65 Save up to $161.54
  • Rent Book $140.79
    Add to Cart Free Shipping Icon Free Shipping

    *This item is part of an exclusive publisher rental program and requires an additional convenience fee. This fee will be reflected in the shopping cart.

Supplemental Materials

What is included with this book?


In the twelve years since publication of Invertebrates, Second Edition, fundamental shifts have occurred in our understanding of the origins and evolutionary relationships among protists and animals. These changes are largely due to the explosion of molecular phylogenetics and evo-devo
research, emergence of the new field of animal genomics, major fossil discoveries in China, Australia, and elsewhere, and important new embryological and ultrastructural studies. As a result:

- New phyla have been described (e.g., Micrognathozoa, Xenacoelomorpha).
- Old phyla have been collapsed into others (e.g., Sipuncula and Echiura are now placed within Annelida; acanthocephalans are now known to be highly modified, parasitic rotifers).
- Phyla once thought to be deuterostomes are now part of the protostome clade (e.g., Chaetognatha, Phoronida, Bryozoa, Brachiopoda).
- The Protostomia has been reorganized into two major clades known as Ecdysozoa and Spiralia.

For each of the thirty-two currently recognized phyla, Invertebrates, Third Edition, presents detailed classifications, revised taxonomic synopses, updated information on general biology and anatomy, and current phylogenetic hypotheses, organized with boxes and tables, and illustrated with abundant
line drawings and new color photos. The chapters are organized around the "new animal phylogeny," while introductory chapters provide basic background information on the general biology of invertebrates. Two new coauthors have been added to the writing team, and twenty-two additional invertebrate
zoologists have contributed to chapter revisions. This benchmark volume on our modern views of invertebrate biology should be in every zoologist's library.

For Instructors

Instructor's Resource Library: Available to qualified adopters, the Instructor's Resource Library for Invertebrates, Third Edition, contains an extensive collection of images for use in teaching the course:

- Textbook Figures and Tables: All of the textbook's figures and tables are included as both high- and low-resolution JPEGs, for easy use in presentation software, learning management systems, and assessments. New for the Third Edition, this now includes all of the textbook's photographs.
- Supplemental Photo Collection: This collection of over 900 photographs depicts organisms that span the entire range of phyla covered in the textbook.
- PowerPoint Presentations: Two ready-to-use PowerPoint presentations are provided for each chapter of the textbook: one that contains all of the textbook figures and tables, and one that contains all of the relevant photos from the supplemental photo collection.

Author Biography

Richard C. Brusca is Executive Director, Emeritus of the Arizona-Sonora Desert Museum and a Research Scientist at the University of Arizona. Rick is the author of nearly 200 research publications and 13 books, including the popular field guides Common Intertidal Invertebrates of the Gulf of California, A Seashore Guide to the Northern Gulf of California, and A Natural History the Santa Catalina Mountains, with an Introduction to the Madrean Sky Islands. He has been the recipient of more than 100 research grants from the National Science Foundation, NOAA, the National Geographic Society, the David and Lucile Packard Foundation, and many other agencies and foundations. He has served on panels for the National Science Board, National Science Foundation, Smithsonian Institution, PEW Program in Conservation and the Environment, Public Broadcasting Service, IUCN Species Survival Commission, and many others. Rick has also served on many environmental non-profit boards, in the U.S. and abroad, and he has organized and conducted field expeditions throughout the world, on every continent. He is an elected Fellow in the American Association for the Advancement of Science (AAAS), the Linnean Society of London (FLS), and the California Academy of Sciences.

Wendy Moore is Assistant Professor in the Department of Entomology at the University of Arizona and Curator of the University of Arizona Insect Collection. Her degrees were earned at Vanderbilt University (B.S., General Biology), the College of Charleston (M.S., Marine Biology), and the University of Arizona (Ph.D., Entomology/Ecology and Evolutionary Biology). Dr. Moore's long-term research interest is the evolution of biotic diversity--especially the evolution of symbiotic lifestyles and how major biotic, climatic, and tectonic events may have influenced the timing and patterns of diversification. Much of her current research is on the carabid beetle subfamily Paussinae, many species of which are obligate symbionts with ants. She is also deeply committed to collections care and enhancement, and the use of bioinformatics to make collections-based data widely available to diverse user communities.

Stephen M. Shuster is Professor of Invertebrate Zoology and Curator of Marine Invertebrates and Molluscs at Northern Arizona University. He earned a B.S. in Zoology from the University of Michigan, an M.S. in Biology from the University of New Mexico, and a Ph.D. in Zoology from the University of California, Berkeley. The author or coauthor of over 75 journal articles, encyclopedia entries, book reviews, and contributed book chapters, Dr. Shuster collaborated with Michael J. Wade on the book Mating Systems and Strategies (2003). His research broadly concerns mating system evolution, male and female reproductive behavior, community and ecosystem genetics, and the population biology of marine organisms. His recent work focuses on the measurement of selection within and among species, and the maintenance of genetic variation in natural populations of marine crustaceans and terrestrial arthropods.

Table of Contents

1. Introduction
Keeping Track of Life
Prokaryotes and Eukaryotes
Where Did Invertebrates Come From?
The Dawn of Life
The Ediacaran Period and the Origin of Animals
The Paleozoic Era (541-252 Ma)
The Mesozoic Era (252-66 Ma)
The Cenozoic Era (66 Ma-present)
Where Do Invertebrates Live?
Marine Habitats
Estuaries and Coastal Wetlands
Freshwater Habitats
Terrestrial Habitats
A Special Type of Environment: Symbiosis
Biodiversity Patterns
New Views of Invertebrate Phylogeny
Some Comments on Evolution
A Final Introductory Message to the Reader

2. Systematics, Phylogeny, and Classification
Biological Classification
Monophyly, Paraphyly, and Polyphyly
Characters and the Concept of Homology
Phylogenetic Trees
Pleisiomorphy and Apomorphy
Constructing Phylogenies and Classifications
Molecular Phylogenetics

3. The Protists: Kingdom Protista
Taxonomic History and Classification
Overviews of the Major Clades, or Groups
The General Protistan Body Plan
The Protist Phyla
Group 1: Amoebozoa
Phylum Amoebozoa: Amebas
Group 2: Chromalveolata
Phylum Dinoflagellata: Dinoflagellates
Phylum Apicomplexa: Gregarines, Coccidians, Haemosporidians, and Their Kin
Phylum Ciliata: The Ciliates
Phylum Stramenopila: Diatoms, Brown Algae, Golden Algae, Slime Nets, Oomycetes, etc.
Phylum Haptophyta: Coccolithophores
Phylum Cryptomonada: Cryptomonads
Group 3: Rhizaria
Phylum Chlorarachniophyta: Chlorarachniophyte Algae
Phylum Granuloreticulosa: Foraminiferans and Their Kin
Phylum Radiolaria: Radiolarians
Phylum Haplosporidia: Haplosporidians
Group 4: Excavata
Phylum Parabasalida: Trichomonads, Hypermastigotes, and Their Kin
Phylum Diplomonadida: Diplomonads
Phylum Heterolobosea: Heterolobosids
Phylum Euglenida: Euglenids
Phylum Kinetoplastida: Trypanosomes, Bodonids, and Their Kin
Group 5: Opisthokonta
Phylum Choanoflagellata: Choanoflagellates
Protist Phylogeny
The Origin of the Protista
Relationships among the Protists

4. Introduction to the Animal Kingdom: Animal Architecture and
Body Plans
Body Symmetry
Cellularity, Body Size, Germ Layers, and Body Cavities
Locomotion and Support
Feeding and Digestion
Excretion and Osmoregulation
Circulation and Gas Exchange
Nervous Systems and Sense Organs
Nervous Systems and Body Plans
Hormones and Pheromones
Asexual Reproduction
Sexual Reproduction

5. Introduction to the Animal Kingdom: Development, Life Histories,
and Origin
Evolutionary Developmental Biology-EvoDevo
Developmental Tool Kits
The Relationship Between Genotype and Phenotype
The Evolution of Novel Gene Function
Gene Regulatory Networks
Eggs and Embryos
Orientation of Cleavage Planes
Radial and Spiral Cleavage
Cell Fates
Blastula Types
Gastrulation and Germ Layer Formation
Mesoderm and Body Cavities
Life Cycles: Sequences and Strategies
Classification of Life Cycles
Indirect Development
Settling and Metamorphosis
Direct Development
Mixed Development
Adaptations to Land and Fresh Water
Parasite Life Cycles
The Relationships Between Ontogeny and Phylogeny
The Origin of the Metazoa

6. Two Basal Metazoan Phyla: Porifera and Placozoa
Phylum Placozoa
Phylum Porifera: The Sponges
Taxonomic History and Classification
The Poriferan Body Plan
Some Additional Aspects of Sponge Biology
Distribution and Ecology
Biochemical Agents
Growth Rates
Poriferan Phylogeny
The Origin of Sponges
Evolution within the Porifera

7. Phylum Cnidaria: Anemones, Corals, Jellyfish, and Their Kin
Taxonomic History and Classification
The Cnidarian Body Plan
Cnidarian Phylogeny
Ediacaran Cnidaria?
Cnidarian Origins
Relationships within Cnidaria

8. Phylum Ctenophora: The Comb Jellies
Taxonomic History and Classification
The Ctenophoran Body Plan
Ctenophoran Phylogeny

9. Introduction to the Bilateria and the Phylum Xenacoelomorpha: Triploblasty and Bilateral Symmetry Provide New Avenues for Animal Radiation
The Basal Bilaterian
Protostomes and Deuterostomes
Phylum Xenacoelomorpha
Classification of Phylum Xenacoelomorph
Class Acoela
The Acoel Body Plan
Class Nemertodermatida
The Nemertodermatid Body Plan
Subphylum Xenoturbellida
The Xenoturbellid Body Plan

10. Phylum Platyhelminthes: The Flatworms
Taxonomic History and Classification
The Platyhelminth Body Plan
Platyhelminth Phylogeny

11. Four Enigmatic Protostome Phyla: Rhombozoa, Orthonectida,
Chaetognatha, Gastrotricha
Phylum Rhombozoa
The Dicyemida
The Heterocyemida
Phylum Orthonectida
Phylum Chaetognatha
Chaetognath Classification
The Chaetognath Body Plan
Phylum Gastrotricha: The Gastrotrichs
Gastrotrich Classification
The Gastrotrich Body Plan

12. Phylum Nemertea: The Ribbon Worms
Taxonomic History and Classification
The Nemertean Body Plan
Nemertean Phylogeny

13. Phylum Mollusca
Taxonomic History and Classification
The Molluscan Body Plan
Molluscan Evolution and Phylogeny

14. Phylum Annelida: The Segmented (and Some Unsegmented)
Taxonomic History and Classification
The Annelid Body Plan
Sipuncula: The Peanut Worms
Sipunculan Classification
The Sipunculan Body Plan
Echiuridae: The Spoon Worms
Siboglinidae: Vent Worms and Their Kin
Siboglinid Taxonomic History
The Siboglinidae Body Plan
Hirudinoidea: Leeches and Their Relatives
The Hirudinoidean Body Plan
Annelid Phylogeny

15. Two Enigmatic Spiralian Phyla: Entoprocta and Cycliophora
Phylum Entoprocta: The Entoprocts
Entoproct Classification
The Entoproct Body Plan
Phylum Cycliophora: The Cycliophorans

16. The Gnathifera: Phyla Gnathostomulida, Rotifera (including
Acanthocephala), and Micrognathozoa
Phylum Gnathostomulida: The Gnathostomulids
Gnathostomulid Classification
The Gnathostomulid Body Plan
Phylum Rotifera: The Free-Living Rotifers
Rotifer Classification
The Rotifer Body Plan
Phylum Rotifera, Subclass Acanthocephala: The Acanthocephalans
The Acanthocephalan Body Plan
Phylum Micrognathozoa: The Micrognathozoans
The Micrognathozoan Body Plan

17. The Lophophorates: Phyla Phoronida, Bryozoa, and Brachiopoda
Taxonomic History of the Lophophorates
The Lophophorate Body Plan
Phylum Phoronida: The Phoronids
The Phoronid Body Plan
Phylum Bryozoa: The Moss Animals
The Bryozoan Body Plan
Phylum Brachiopoda: The Lamp Shells
The Brachiopod Body Plan

19. The Nematoida: Phyla Nematoda and Nematomorpha
Phylum Nematoda: Roundworms and Threadworms
Nematode Classification
The Nematode Body Plan
Life Cycles of Some Parasitic Nematodes
Phylum Nematomorpha: Horsehair Worms and Their Kin
The Nematomorphan Body Plan

19. The Scalidophora: Phyla Kinorhyncha, Priapula, and Loricifera
Phylum Kinorhyncha: The Kinorhynchs
Kinorhynch Classification
The Kinorhynch Body Plan
Phylum Priapula: The Priapulans
Priapulan Classification
The Priapulan Body Plan
Phylum Loricifera: The Loriciferans

20. The Emergence of the Arthropods: Tardigrades, Onychophorans,
and the Arthropod Body Plan
Phylum Tardigrada
The Tardigrade Body Plan
Phylum Onychophora
The Onychophoran Body Plan
An Introduction to the Arthropods
Taxonomic History and Classification
The Arthropod Body Plan and Arthropodization
The Evolution of Arthropods
The Origin of Arthropods
Evolution within the Arthropoda

21. Phylum Arthropoda: Crustacea: Crabs, Shrimps, and Their Kin
Classification of The Crustacea
Synopses of Crustacean Taxa
The Crustacean Body Plan
Crustacean Phylogeny

22. Phylum Arthropoda: The Hexapoda: Insects and Their Kin
Hexapod Classification
The Hexapod Body Plan
Hexapod Evolution

23. Phylum Arthropoda: The Myriapods: Centipedes, Millipedes,
and Their Kin
Myriapod Classification
The Myriapod Body Plan
Myriapod Phylogeny

24. Phylum Arthropoda: The Chelicerata
Chelicerate Classification
The Euchelicerate Body Plan
The Class Pycnogonida
The Pycnogonid Body Plan
Chelicerate Phylogeny

25. Introduction to the Deuterostomes and the Phylum
Phylum Echinodermata
Taxonomic History and Classification
The Echinoderm Body Plan
Echinoderm Phylogeny
First Echinoderms
Modern Echinoderms

26. Phylum Hemichordata: Acorn Worms and Pterobranchs
Hemichordate Classification
The Hemichordate Body Plan
Enteropneusta (Acorn Worms)
Hemichordate Fossil Record and Phylogeny

27. Phylum Chordata: Cephalochordata and Urochordata
Chordate Classification
Phylum Chordata, Subphylum Cephalochordata: The Lancelets (Amphioxus)
The Cephalochordate Body Plan
Phylum Chordata, Subphylum Urochordata: The Tunicates
The Tunicate Body Plan
Chordate Phylogeny

28. Perspectives on Invertebrate Phylogeny

Illustration Credits

Supplemental Materials

What is included with this book?

The New copy of this book will include any supplemental materials advertised. Please check the title of the book to determine if it should include any access cards, study guides, lab manuals, CDs, etc.

The Used, Rental and eBook copies of this book are not guaranteed to include any supplemental materials. Typically, only the book itself is included. This is true even if the title states it includes any access cards, study guides, lab manuals, CDs, etc.

Rewards Program